Education

- 1985-1988: B.Sc. with Honours (Class I) in Biochemistry, Department of Biochemistry, University of Leeds, Leeds, U.K..
- 1988-1992: Ph.D. in Biology/Cytology, Department of Biology (supervisors : N. Avrion Mitchison and Peter C.L.Beverley), University College, University of London, London, U.K.. Thesis on the "Control of Transcription in Embryonal Carcinoma Cells", conducted in the Laboratory of Eukaryotic Molecular Genetics (supervisor: Nicholas B. La Thangue), MRC National Institute for Medical Research, Mill Hill, London, U.K..
- 1998-2000: Post-graduate Certificate in Open and Distance Education, Greek Open University, Patra, Greece.